1 | #Please insert up references in the next lines (line starts with keyword UP) |
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2 | UP arb.hlp |
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3 | UP glossary.hlp |
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4 | |
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5 | #Please insert subtopic references (line starts with keyword SUB) |
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6 | SUB mode_setroot.hlp |
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7 | SUB syncroots.hlp |
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8 | SUB mode_group.hlp |
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9 | |
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10 | # Hypertext links in helptext can be added like this: LINK{ref.hlp|http://add|bla@domain} |
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11 | |
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12 | #************* Title of helpfile !! and start of real helpfile ******** |
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13 | TITLE Keeled groups |
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14 | |
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15 | DESCRIPTION Taxonomic groups in an unrooted tree(!) can be defined by |
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16 | * the branch at which the group is located |
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17 | * the direction into which the group points (i.e. which of |
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18 | the 2 subtrees that are connected by this branch does form |
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19 | the defined group) |
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20 | |
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21 | In rooted trees the branch location on its own suffices as |
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22 | group-defining criteria, if we assume all groups point away from |
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23 | the root. This is the way taxonomic groups are handled in most |
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24 | software and so did arb (up to version 6.0.6). |
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25 | |
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26 | This simple handling of groups was not able to reflect the semantic change |
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27 | which is implicitely applied by moving the trees root into a group. |
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28 | |
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29 | To solve this problem, arb-7.0 introduced the concept of "keeled groups". |
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30 | |
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31 | A keeled group may be interpreted in 2 (semantically equivalent) ways: |
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32 | * the keeled group "points upwards" (i.e. semantically it defines a taxonomic unit containing the two OTHER subtrees starting at its parent-node) |
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33 | * the keeled group "graphically embraces the inverse subset of species" compared with the original unkeeled group |
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34 | |
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35 | To indicate keeled groups, their names are prefixed by exclamation |
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36 | marks, e.g. shown as '!group' which should be read as "not group"; |
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37 | this emphasizes the 2nd interpretation given above. |
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38 | |
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39 | Example: |
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40 | |
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41 | When the root of the tree is placed somewhere inside a |
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42 | group named 'Bacteria', that group gets keeled over. |
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43 | Its name now gets displayed as '!Bacteria' (should read |
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44 | as "not Bacteria") and the group does embrace the rest |
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45 | of the tree, i.e. "everything but Bacteria". |
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46 | |
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47 | |
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48 | NOTES Two groups with different orientation (i.e. a normal and a |
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49 | keeled group) may appear at the same inner node. |
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50 | This happens e.g. when the original tree had groups defined |
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51 | at both subtrees starting directly at root. |
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52 | The group name of such a group will be "Normal = !Keeled". |
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53 | |
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54 | SECTION Howto 'fix' keeled groups |
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55 | |
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56 | It's not possible to export keeled groups into newick files! |
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57 | |
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58 | If a tree contains keeled groups, there is a mismatch between |
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59 | - the taxonomy defined by the groups in the tree and |
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60 | - the position of the tree-root. |
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61 | |
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62 | One reason for that mismatch is a misplaced tree-root. |
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63 | In that case there exists at least one root-position where no |
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64 | groups are keeled. |
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65 | To find such a position use LINK{mode_setroot.hlp} and |
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66 | repeatedly click on any keeled group you spot, |
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67 | until they've all disappeared. |
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68 | |
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69 | If they won't disappear, the taxonomy of the tree probably is |
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70 | not well-defined and has to be resolved manually. |
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71 | In that case the following tools may help: |
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72 | |
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73 | - right click onto a keeled group in LINK{mode_group.hlp} to |
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74 | toggle its keeled state, |
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75 | - afterwards use LINK{mode_move.hlp} to correct the position |
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76 | of that group OR |
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77 | - delete a keeled group, if it does no longer fit into the tree. |
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78 | This might e.g. apply after topology changes. |
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79 | |
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80 | EXAMPLES None |
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81 | |
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82 | WARNINGS None |
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83 | |
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84 | BUGS No bugs known |
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